The Laboratory for Molecular Cell Biology (LMCB) at UCL is dedicated to research aimed at understanding the molecular and cellular basis of cell behaviour relevant to human disease. This four-year Molecular Cell Biology PhD programme at the Laboratory for Molecular Cell Biology (LMCB) provides a broad in-depth training in molecular cell biology and fundamental aspects of biomedical science. The programme provides students with the opportunity to carry out cutting-edge research in an internationally renown environment. First-year rotations through three different laboratories and a bespoke tutorial series just are some of the programme highlights. The goals of our training are to produce scientists who have explored one area and one level of analysis in great depth, but who are familiar with the full scope of molecular and cell biology. They should be able to move from one level to another in a critical and creative manner. We also try to develop an appreciation for translational research that bears on human disease. During their first year, students have a free choice of three laboratories from the groups within the LMCB, and spend three months in each, working on research projects.